CAMB 5110

Principles of Development

University of Pennsylvania · UGRD · Fall 2026

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This graduate course, which will include lectures and readings from the literature, is designed to provide a foundation in the principles of developmental biology. Topics covered will include: the germ line and piRNA, signaling pathways in development, pattern formation and cell specification, gastrulation, tissue differentiation, morphogenesis, cell polarity, epigenetics in development, organogenesis, stem cell biology, regeneration, and developmental evolution. The use of molecular biology, biochemistry, genomics and genetics, cell biology, and embryological manipulations will be discussed in the context of the analysis of developmental mechanisms. Undergraduate background in cell biology and molecular biology required. Non-BGS students require permission from course director to enroll.

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Class #pennsylvania_2-CAMB5110Fall 2026UGRD1 credits
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